Takagi, S., Hosaka, T., & Okuda, T. 2005 Material of dipterocarp-associated gall-inhabiting coccoids collected in Negeri Sembilan, Malaya (Homoptera: Coccoidea).. Insecta Matsumurana (New Series) 62: 123-151.

Notes: Four species of dipterocarp-associated gall-inhabiting coccoids were collected in Negeri Sembilan, Malaya, Malaysia, in 2003 and 2004 in surveys conducted under a project on biodiversity in tropical forests, and taxonomic accounts of them are given in this paper. One of them, Gallacoccus longisetosus, sp. nov., is described on the basis of two female and three male instars. This species is probably rightly referred to Gallacoccus, and yet peculiar among the known species of the genus in having spinous but remarkably elongated setae dorsally on the first-instar female. The other species are also referred to Gallacoccus, but they are not formally named owing to the insufficient material; instead, they are designated by tags coined by combining abbreviations of the collection year, locality, and host plant. One of them is closely similar to G. longisetosus in many adult male characters, whereas the other two resemble Gallacoccos heckrothi in the male nymphs or the adult female. Some features and their characters are discussed with regard to their significance in function and phylogeny: dermal pits in the first-instar female, legs in the first instar male, penial sheath pores and other sensilla, and papillae and claws in the adult male. The discussions remain preliminary, but suggest intricate relationships between function and phylogeny.