Valid Names Results

Madarococcus intermedius (Maskell, 1891) (Eriococcidae: Madarococcus)

Nomenclatural History

• Rhizococcus intermedius Maskell 1891: 19-20. Type data: NEW ZEALAND: South Island, Reefton, on Fagus menziesii, ?/03/1890, by W.M. Maskell. Lectotype, female, by subsequent designation (DeitzTo1980,47). Type depository: Auckland: New Zealand Arthropod Collection, Landcare Research, New Zealand; accepted valid name Notes: Type material also in USNM Illustr.
• Nidularia intermedius (Maskell, 1891); Lindinger 1933a: 116. change of combination
• Sisyrococcus intermedius (Maskell, 1891); Hoy 1962: 182. change of combination Illustr.
• Madarococcus intermedius (Maskell, 1891); Hardy, Beardsley & Gullan 2008: 383-384. change of combination Notes: In addition to the lectotype, the NZAC has 1, three original slides: one labelled 'larva'; one labelled 'female 2nd-stage' [actually an adult female]; one labelled '2 males', and 2, 16 slides mounted by Hoy, 14 labelled 'Mask Coll. / Ex Nothofagus menziesii leaf axils /1890 [no coll no.],' and two with same data except for a more specific date, March 1890. (Hardy, et al., 2009)

Common Names

• beech rhizococcus Miller1925

Ecological Associates

Hosts:

Families: 1 | Genera: 1

Geographic Distribution

Keys

• HardyGuHo2008: pp.375-376 ( Adult (F) ) [Key to species of Madarococcus based on adult females]
• Hoy1962: pp.181 ( Adult (F) ) [Sisyrococcus species of New Zealand] Key as: Sisyrococcus intermedius

Remarks

• Systematics: Slide-mounted adult female with: enlarged setae conical, sides concave basally, straight elsewhere, apices rounded, marginal setae and medial setae on abdomen conspicuously longer than all other dorsal setae, bases of many setae with 1 associated microtubular duct; body shape unusual, anterior abdominal segments overlying posterior segments; multilocular pores in cluster on dorsum of head; macrotubular ducts confined to last 5 abdominal segments; microtubular ducts elongate, without sclerotized area (Hoy, 1962). Hoy’s two species of Sisyrococcus, now M. intermedius and M. papillosus, have very unusual adult females. In mature specimens, the dorsal surface of the anterior abdominal segments is folded back and over the posterior segments, concealing the anal lobes from the dorsal aspect. Macrotubular ducts are found only on the dorsal surface of the abdomen and, on the anterior segments, they are arranged in tight, ransverse rows. (Hardy, et al., 2008) Madarococcus intermedius can be distinguished from M. papillosus by having far fewer dorsal setae, and all but a few pairs confined to the margin (in M. papillosus the dorsum is covered with c. 500 setae and the margin is indistinguishable). M. intermedius is unique among Madarococcus species in having quinquelocular disc pores on the dorsum, in a narrow medial band on the head. The suranal setae on M. intermedius and M. papillosus often appear non-spatulate, and in young adult females they can have fine flagellate tips that later break off. (Hardy, et al., 2008)
• Structure: This species apparently has no sac. Glassy filaments arise from enlarged setae of margin and dorsum (Hoy, 1962). Adult female sub-globular, naked, sometimes conspicuously segmented, sometimes nearly smooth. Female is red, yellow or greenish in color. First instars reddish-yellow, active, naked, flattish, elliptical, tapering posteriorly. Male pupae in a dull white closely felted sac. Adult male is brown (Maskell, 1891). Tests of males are creamy white and occur on the undersides of leaves, especially near the leaf margins. (Hardy, et al., 2008)
• Biology: Mature females shelter embryos in a cavity under their body. (Hardy, et al., 2008)
• General Remarks: Detailed descriptions and illustrations by Maskell (1891) and Hoy (1962).

Illustrations

Citations