Valid Names Results
Dysmicoccus boninsis (Kuwana, 1909) (Pseudococcidae: Dysmicoccus)Nomenclatural History
- Dactylopius calceolariae; Maskell 1890: 149. misidentification
- Pseudococcus calceolariae; Kirkaldy 1902: 103. misidentification
- Trechocorys calceolariae; Kirkaldy 1904: 227. misidentification
- Dactylopius (Pseudococcus) boninsis Kuwana 1909a: 161. Type data: JAPAN: Ogasawara (Bonin) Islands, on sugar cane. Syntypes, female, accepted valid name Notes: Type material probably lost, see Zimmerman (1948) and Beardsley (1966a).
- Pseudococcus boninensis (Kuwana, 1909); Lindinger 1914: 158. emendation that is unjustified
- Dactylopius boninsis; Kuwana 1917a: 169. subsequent use
- Pseudococcus heterospinus Green 1923d: 91. Type data: MADEIRA: Machico, on grasses. Lectotype, female, by subsequent designation (Willia1985,116). Type depository: London: The Natural History Museum, England, UK; junior synonym (discovered by Balach1939, 260). Illustr.
- Trionymus calceolariae; Fullaway 1923: 308. misidentification
- Pseudococcus boninsis (Kuwana, 1909); Morrison 1925a: 487. change of combination
- Pseudococcus aegyptiacus Hall 1925: 8. Type data: EGYPT: Mit Ghamr, on Saccharum officinarum. Holotype, female, Type depository: London: The Natural History Museum, England, UK; junior synonym (discovered by Hall1926a, 33). Illustr.
- Trionymus boninsis (Kuwana, 1909); Swezey 1931: 946. change of combination
- Trionymus taiwanus Takahashi 1932: 41. Type data: TAIWAN: Taihoku, on Oryza sativa. Syntypes, female, Type depository: Taichung: Entomology Collection, Taiwan Agricultural Research Institute, Wu-feng, Taichung, Taiwan; junior synonym (discovered by Willia1970DJ, 128).
- Erium boninense (Kuwana, 1909); Lindinger 1935a: 122. change of combination
- Erium taiwanum (Takahashi, 1932); Lindinger 1935a: 122. change of combination
- Erium boninense (Kuwana, 1909); Lindinger 1935a: 122. emendation that is unjustified
- Pseudococcus zeae Kanda 1943: 49. Type data: JAPAN: Mitsuzawa, Yokohama, on Zea mays and on Syntherisma sanguinalis var. ciliaris. Syntypes, female, Type depository: Bucarest: Academie des Sciences Agricoles et Forestieres, Romania; junior synonym (discovered by Kawai1980, 104).
- Dysmicoccus boninsis (Kuwana, 1909); Ferris 1950b: 57. change of combination
- Trionymus kayashimai Takahashi 1951a: 14. Type data: MALAYSIA: West Malaysia, Selangor, Gombak, on Saccharum arundinaceum, 30.v.1944, collected R. Takahashi.. Lectotype, female, by subsequent designation (Willia1999,92). Type depository: London: The Natural History Museum, England, UK; junior synonym (discovered by Willia1999, 92).
- Trionymus merrilli Ferris 1953a: 488. Type data: U.S.A.: Florida, Fort Pierce, on 'Para Grass'. Syntypes, female, Type depository: Davis: The Bohart Museum of Entomology, University of California, California, USA; junior synonym (discovered by MillerMi2002a, 969-970). Illustr.
- Dysmicoccus merrilli (Ferris, 1953); McKenzie 1960: 706. change of combination
- Vryburgia graminea De Lotto 1967: 25. Type data: SOUTH AFRICA: Durban, on grass. Holotype, female, Type depository: Pretoria: South African National Collection of Insects, South Africa; junior synonym (discovered by DeLott1977, 29).
- Dysmicoccus boniansis (Kuwana, 1909); Ali 1970a: 105. misspelling of species epithet
- Trionymus boninsis (Kuwana, 1909); Danzig & Gavrilov-Zimin 2015: 112. revived combination (previously published)
- Dysmicoccus boninsis (Kuwana, 1909); Williams, D.J. 2017a: 247. revived combination (previously published)
Common Names
- gray sugarcane mealybug Bartle1978e Zimmer1948
- Cochinilla de la caña de azúcar SirisenSaKo2022
- ogasawara-kona-kaigaramushi SirisenSaKo2022
- piojo gris SirisenSaKo2022
- piojo harinoso gris de la caña de azúcar SirisenSaKo2022
Ecological Associates
Hosts:
Families: 12 | Genera: 32
- Araceae
- Araceae | BenDov1994 WilliaGr1992
- Arecaceae
- Cocos nucifera | BenDov1994 WilliaGr1992 WilliaWa1988a
- Asteraceae
- Asteraceae | Moghad2013
- Lactuca | Moghad2013a
- Cannaceae
- Canna | BenDov1994 WilliaGr1992
- Convolvulaceae
- Ipomoea | BenDov1994 WilliaWa1988a
- Cupressaceae
- Juniperus | BenDov1994 WilliaGr1992
- Iridaceae
- Iris | BenDov1994 WilliaGr1992
- Sisyrinchium | BenDov1994 HodgsoHi1990
- Pandanaceae
- Pandanus | BenDov1994 WilliaWa1988a
- Poaceae
- Brachiaria mutica | BenDov1994 WilliaWa1988a BenDov1994 WilliaGr1992 | (= Panicum barbinode)
- Coix lacryma-jobi | Beards1966 BenDov1994 Mamet1949
- Cortaderia | BenDov1994 HodgsoHi1990
- Cortaderia selloana | BenDov1994 WilliaGr1992 | (= Cortaderia argentea)(= Cortaderia argentata)
- Cymbopogon citratus | BenDov1994 WilliaGr1992
- Cynodon | BenDov1994 HodgsoHi1991
- Cynodon dactylon | BenDov1994 WilliaGr1992
- Digitaria ciliaris | BenDov1994 Kanda1943a | (= Syntherisma sanguinalis var. ciliaris)
- Echinochloa | BenDov1994 HodgsoHi1990
- Echinochloa colona | BenDov1994 Tang1992 Willia1970DJ WilliaGr1992
- Eragrostis variabilis | BenDov1994 Zimmer1948
- Gynerium sagittatum | BenDov1994 WilliaGr1992
- Lasiacis divaricata | BenDov1994 WilliaGr1992
- Miscanthus | BenDov1994 KawaiMaUm1971
- Oryza sativa | BenDov1994 LiuTa1988 Willia1970DJ
- Panicum | BenDov1994 HodgsoHi1990 Willia1970DJ
- Panicum maximum | BenDov1994 WilliaWa1988a
- Paspalum | BenDov1994 HodgsoHi1990 KawaiMaUm1971
- Paspalum conjugatum | LincanHoCa2010
- Paspalum distichum | BenDov1994 WilliaGr1992
- Poaceae | BenDov1994 Ferris1953a KawaiMaUm1971 Takaha1941b Willia1970DJ
- Saccharum | BenDov1994 HodgsoHi1990 Kirkal1902
- Saccharum arundinaceum | BenDov1994 Takaha1951a
- Saccharum officinarum | BenDov1994 Granar1991 KawaiMaUm1971 Laing1933 Takaha1932 WilliaGr1992 WilliaWa1988a
- Saccharum spontaneum | Willia2004a
- Sorghum | BenDov1994 WilliaGr1992
- Sorghum arundinaceum | BenDov1994 Tang1992 Willia1985 | (= Sorghum verticilliflorum)
- Sorghum halepense | BenDov1994 WilliaGr1992 WilliaWa1988a
- Stenotaphrum | BenDov1994 HodgsoHi1990
- Stenotaphrum secundatum | BenDov1994 WilliaGr1992
- Thysanolaena latifolia | BenDov1994 Mamet1949 | (= Thysanolaena agrostis)
- Tripsacum | BenDov1994 WilliaGr1992
- Zea mays | BenDov1994 Kanda1943a Willia1970DJ WilliaGr1992
- Rutaceae
- Citrus | BenDov1994 Marott1987a
- Solanaceae
- Solanum lycopersicum | PedrazVaGo2019
- Vitaceae
- Cissus alata | BenDov1994 Marott1987a | (= Cissus rhombifolia)
Foes:
Families: 1 | Genera: 3
- Encyrtidae
- Pseudaphycus mundus | Moore1988
- Pseudococcobius terryi | Bynum1937
- Rhopus nigroclavatus | AbdRab2001d
Associates:
Families: 2 | Genera: 9
- Betaproteobacteria
- Candidatus Tremblaya princeps | ChoiLe2022
- Formicidae
- Brachymyrmex | KondoRaVe2008
- Crematogaster | KondoRaVe2008
- Ectatomma ruidum | KondoRaVe2008
- Linepithema | KondoRaVe2008
- Nylanderia | KondoRaVe2008
- Pheidole | KondoRaVe2008
- Pheidole biconstricta | CaballRaKo2017
- Solenopsis | KondoRaVe2008
- Wasmannia auropunctata | KondoRaVe2008
Geographic Distribution
Countries: 63
- Afghanistan | KozarFoZa1996
- Antigua and Barbuda
- Antigua | BenDov1994 Willia1970DJ WilliaGr1992
- Barbuda | BenDov1994 CABI1960 WilliaGr1992
- Argentina
- Tucuman | BenDov1994 Ferris1950b Granar1991 Granar2009 WilliaGr1992
- Australia
- Queensland | BenDov1994 Willia1985
- Bahamas | BenDov1994 Willia1970DJ WilliaGr1992
- Barbados | SirisenSaKo2022
- Bermuda | BenDov1994 Ferris1950b HodgsoHi1990 HodgsoHi1991 Willia1970DJ WilliaGr1992
- Bonin Islands (=Ogasawara-Gunto) | Ali1970a BenDov1994 Ferris1950b Kawai1987 KawaiMaUm1971 Kuwana1909a | SirisenSaKo2022
- Brazil
- Bahia | SirisenSaKo2022
- Espirito Santo | CulikGu2005 CulikMaVe2007
- Minas Gerais | SirisenSaKo2022
- Pernambuco | SirisenSaKo2022
- Rio de Janeiro | SirisenSaKo2022
- Sao Paulo | BenDov1994 CABI1960 Ferris1950b WilliaGr1992
- Cape Verde | SchmutPiKl1978 VanHarCoWi1990
- China | BenDov1994 Tang1992 TangHaTa1992 Wu1935
- Colombia | BenDov1994 Kondo2001 KondoRaVe2008 WilliaGr1992
- Cote d'Ivoire (=Ivory Coast) | BenDov1994 CouturMaRi1985
- Cuba | BenDov1994 CABI1960 WilliaGr1992 WilliaMaSu2001
- Dominica | SirisenSaKo2022
- Dominican Republic | BenDov1994 CABI1960 Ferris1950b Schmut1990 WilliaGr1992
- Ecuador | BenDov1994 WilliaGr1992
- Egypt | BenDov1994 Hall1925 Willia1970DJ
- Federated States of Micronesia
- Caroline Islands | Beards1966 BenDov1994
- Pohnpei Island | Beards1966 BenDov1994
- Truk Islands | Beards1966 BenDov1994
- Fiji | BenDov1994 HodgsoLa2011 Maskel1890 WilliaWa1988a
- Galapagos Islands | CaustoPeSi2006 LincanHoCa2010
- Grenada | BenDov1994 Willia1970DJ WilliaGr1992
- Guadeloupe | Balach1957c BenDov1994 CABI1960 MatileEt2006
- Guam | Beards1966 BenDov1994
- Haiti | BenDov1994 CABI1960 PerezG2008 WilliaGr1992
- Hawaiian Islands
- India | SirisenSaKo2022
- Indonesia
- Sumatra | Willia2004a
- Iran | KozarFoZa1996 Moghad2009
- Italy
- Sicily | BenDov1994 Marott1987a
- Jamaica | BenDov1994 Kirkal1902 Willia1970DJ WilliaGr1992
- Japan | BenDov1994 Ferris1950b Kawai1980 KawaiMaUm1971
- Madeira Islands | BenDov1994 Willia1970DJ
- Malaysia | BenDov1994 Takaha1951a Willia1999 Willia2004a
- Martinique | Balach1957c BenDov1994 MatileEt2006
- Mexico | BenDov1994 CABI1960 WilliaGr1992
- Montserrat | BenDov1994 WilliaGr1992
- Mozambique | BenDov1994 Saraiv1939
- New Britain | BenDov1994 CABI1960 | SirisenSaKo2022
- New Caledonia | BenDov1994 Laing1933 Willia1970DJ WilliaWa1988a
- New Zealand | BenDov1994 Kirkal1902
- Niue | BenDov1994 WilliaWa1988a
- Northern Mariana Islands | BenDov1994 CABI1960
- Saipan Island | BenDov1994 Takaha1941b
- Palau | Beards1966 BenDov1994
- Panama | BenDov1994 CABI1960 Ferris1950b WilliaGr1992
- Puerto Rico & Vieques Island
- Puerto Rico | BenDov1994 Martor1976 Willia1970DJ WilliaGr1992
- Vieques Island | BenDov1994 CABI1960
- Reunion | GermaiMiPa2014 Mamet1943a
- Rodriques Island | BenDov1994 CABI1960 Mamet1949
- Ryukyu Islands (=Nansei Shoto) | BenDov1994 Takaha1940a | SirisenSaKo2022
- Singapore | Willia2004a
- South Africa | BenDov1994 DeLott1967 DeLott1977
- Spain | BeltraSo2011
- Sri Lanka | Willia2004a
- Suriname | BenDov1994 Willia1970DJ WilliaGr1992
- Taiwan | Ali1970a BenDov1994 LiuTa1988 Takaha1932
- Tonga | BenDov1994 WilliaWa1988a
- Trinidad and Tobago
- Trinidad | BenDov1994 Willia1970DJ WilliaGr1992
- Tromelin Island | BenDov1994 Mamet1952
- Tuvalu | BenDov1994 WilliaWa1988a
- U.S. Virgin Islands | BenDov1994 CABI1960 WilliaGr1992
- United States
- Florida | BenDov1994 DownieGu2004 Ferris1950b Willia1970DJ
- Georgia | BenDov1994 Ferris1950b
- Louisiana | BenDov1994 Ferris1950b
- Mississippi | BenDov1994 Willia1970DJ
- Venezuela | BenDov1994 Willia1970DJ WilliaGr1992
- Western Samoa | BenDov1994 Willia1970DJ WilliaWa1988a | SirisenSaKo2022
Keys
- ZhangWaWa2023: pp.129 ( Adult (F) ) [Dysmicoccus species in China]
- ZarkanErAp2023: pp.164-165 ( Adult (F) ) [Dysmicoccus in Indonesia]
- EvansAb2023: pp.8-13 ( Adult (F) ) [Pseudococccidae of Egypt]
- MoghadWa2022: pp.42 ( Adult (F) ) [Dysmicoccus in Iran]
- TanakaSaCh2022: pp.309 ( Adult (F) ) [Dysmicoccus species in Japan]
- TanakaKa2021: pp.577-578 ( Adult (F) ) [Dysmicoccus in Japan]
- Hodgso2020: pp.32-34 ( Adult (M) ) [higher groups of Pseudococcidae]
- Moghad2018: pp.185 ( Adult (F) ) [Dysmicoccus species in Iran]
- CaballRaKo2017: pp.498 ( Adult (F) ) [Coccomorpha of sugarcane from Colombia] Key as: Dysmicoccus boninsis
- DanzigGa2015: pp.99-106 ( Adult (F) ) [Palaearctic Trionymus species]
- TanakaUe2012: pp.414 ( Adult (F) ) [Key to separate four new invasive mealybug species: Dysmicoccus neobrevipes, Ph. defectus, Ph. parvus and P. solenopsis from other mealybugs occurring in the Ryukyu Archipelago] Key as: Dysmicoccus boninsis
- Granar2009: pp.12-14 ( Adult (F) ) [Dysmicoccus species of Neotropical region]
- WilliaMa2009a: pp.97-101 ( Adult (F) ) [Pseudococcidae species from Mauritius]
- Willia2004a: pp.161-163 ( Adult (F) ) [Dysmicoccus species of southern Asia] Key as: Dysmicoccus boninsis
- MillerMi2002a: pp.971 ( Adult (F) ) [Key to Species of Dysmicoccus, Oracella and Paradoxococcus of the Gulf States Region (Adult Remales)] Key as: Dysmicoccus boninsis
- Tang1992: pp.151, 192 ( Adult (F) ) [Palaearctic & Oriental regions]
- WilliaWa1988a: pp.46 ( Adult (F) ) [Tropical South Pacific Region]
- Willia1985: pp.108 ( Adult (F) ) [Australia]
- Wang1982ZQ: pp.37 ( Adult (F) ) [China]
- Yang1982: pp.84 ( Adult (F) ) [China]
- Kawai1980: pp.102 ( Adult (F) ) [World]
- DeLott1977: pp.29 ( Adult (F) ) [South Africa]
- DeLott1977: pp.29 ( Adult (F) ) [South Africa] Key as: Dysmicoccus boninsis
- MillerMc1973: pp.504 ( Adult (F) ) [North America] Key as: Dysmicoccus boninsis
- Willia1970DJ: pp.115 ( Adult (F) ) [World]
- McKenz1967: pp.157 ( Adult (F) ) [U.S.A., California] Key as: Dysmicoccus boninsis
- DeLott1967: pp.22 ( Adult (F) ) [South Africa] Key as: Vryburgia graminea
- Beards1966: pp.408 ( Adult (F) ) [Federated States of Micronesia]
- McKenz1964: pp.230 ( Adult (F) ) [North America]
- McKenz1962: pp.644 ( Adult (F) ) [North America]
- Beards1960: pp.212 ( Adult (M) ) [Hawaii]
- Ezzat1960b: pp.40 ( Adult (F) ) [Egypt]
- Ferris1953a: pp.332,483 ( Adult (F) ) [North America]
- Ferris1950b: pp.54 ( Adult (F) ) [North America] Key as: Dysmicoccus boninsis
- Zimmer1948: pp.177 ( Adult (F) ) [Hawaii]
Remarks
- Systematics: Beardsley (1964) reported the occurrence of gray and white-coloured forms, which are taxonomically identical. Prior to 1970 it has frequently been misidentified as Pseudococcus calceolariae (Maskell), Saccharicoccus sacchari (Cockerell) and Pseudococcus saccharifolii (Green) (see Bartlett, 1978e).
- Structure: Dorsum: numerous oral collar tubular ducts present; with only 8 or 9 pairs of cerarii, each containing 2 enlarged setae. Venter: multilocular disc pores present on the fifth and posterior abdominal segments; translucent pores present on hind coxa and a few on the hind tibia; circulus normally notched at each side divided by an intersegmental line, but sometimes small and oval (Williams & Granara de Willink 1992). Slide-mounted adult female oval to broadly oval. Anal lobes quite well developed. Antennae each with 8 segments. Legs well developed; hind leg with translucent pores on dorsal surface of coxa and a few on tibia. (Moghaddam & Watson, 2022).
- Biology: Occurring beneath the leaf sheaths of sugarcane. The life cycle in Hawaii requires from 18 to 26 days (Zimmerman, 1948).
- Economic Importance: The gray sugarcane mealybug has been recorded from a wide range of grasses, as well as hosts from other families, but reported as a pest of sugarcane. Generally it is a minor pest, but when attended by ants it may kill young shoots (Bartlett, 1978e). The encyrtid Aphycus terryi (Fullaway) was found an effective parasitoid in Hawaii, from where it was introduced to Southern USA (Bartlett, 1978e). Beardsley (1960a) noted that it is common on sugarcane in Hawaii and discussed it dispersal, infestation sites and the effect of ants and other factors on its populations.
- General Remarks: Description and illustration of adult female by Kanda (1943), Ferris in Zimmerman (1948), Ferris (1950b), De Lotto (1967) (as Vryburgia graminea), Williams (1970DJ, 1985), Williams & Watson (1988a), Liu & Tao (1988), Williams & Granara de Willink (1992), Williams (2004a) and by Granara de Willink (2009). Description and illustration of adult male by Beardsley (1960).Good description and illustration of the adult female given by Ferris (1953a).
Illustrations
Citations
- AbdRab2001d: biological control, distribution, 1357
- AbdRabEv2021: distribution, host, 41
- Ali1970a: catalog, description, distribution, host, taxonomy, 105
- Balach1939: taxonomy, 260
- Balach1957c: distribution, host, 207
- Bartle1978e: biological control, distribution, economic importance, host, 140, 141
- Beards1960: description, distribution, host, illustration, taxonomy, 216-217
- Beards1960a: biological control, ecology, economic importance, life history, 954-961
- Beards1964: distribution, host, taxonomy, 338
- Beards1966: distribution, host, taxonomy, 408-409
- BeltraSo2011: distribution, host,
- BeltraSoMa2012: molecular data, 167-171
- BenDov1994: catalog, 122,123,135
- Bynum1937: biological control, distribution, host, 756-761
- CaballRaKo2017: diagnosis, distribution, economic importance, host, key, 496
- CaballRaRa2020: host, 169
- CaustoPeSi2006: distribution, 138
- Charpe1956: chemical control, distribution, host, 418-419
- ChenShCh2024: molecular biology, 3
- ChenWoWu2012: distribution, taxonomy, 300
- ChoiLe2022: endosymbiont, phylogeny, 358,359
- CouturMaRi1985: distribution, host,
- CulikGu2005: distribution, host, 1-8
- CulikMaVe2007: distribution, host, 1-5
- DanzigGa2015: distribution, host, taxonomy, 112
- DeLott1967: description, distribution, host, illustration, taxonomy, 25-26
- DeLott1977: distribution, host, taxonomy, 29
- DownieGu2004: distribution, host, molecular data, phylogeny, 258-259
- EvansAb2023: distribution, host, illustration, key, taxonomy, 7, 10, 15, 41
- EzzatNa1987: distribution, 88
- Ferris1950b: description, distribution, host, illustration, taxonomy, 57-58
- Ferris1953a: description, distribution, host, illustration, taxonomy, 488-489
- Flande1943: biological control, 117,128,137
- Flande1951b: biological control, 93-98
- FrancoRuMa2011: distribution, 18,25
- Fullaw1923: 308
- Fullaw1946: distribution, host, taxonomy, 157
- Gavril2018: reproduction, 227
- GermaiMiPa2014: distribution, 24
- Granar1991: distribution, economic importance, host, 261, 262
- Granar2009: description, distribution, host, illustration, taxonomy, 21-22
- Greath1971: biological control, distribution, host,
- Green1923b: 91
- HakkonPi1984: biological control, 1109-1121
- Hall1925: description, distribution, host, illustration, taxonomy, 8, 9
- Hall1926a: taxonomy, 33
- HardyGuHo2008: molecular data, phylogeny, taxonomy, 51-71
- Hodgso2020: key, 33
- Hodgso2020: key, 33
- HodgsoHi1990: distribution, host, 7, 8, 9, 12, 16, 19,
- HodgsoHi1991: distribution, host, 143
- HodgsoLa2011: distribution, host, 27
- Kanda1943: description, distribution, host, illustration, taxonomy, 49-51
- Kawai1980: description, distribution, host, taxonomy, 104
- Kawai1987: distribution, host, 77
- KawaiMaUm1971: distribution, host, taxonomy, 14, 15
- Kirkal1902: distribution, host, 103
- Kirkal1904: biological importance, economic importance, host, taxonomy, 227
- Kondo2001: distribution, host, taxonomy, 35
- KondoRaVe2008: distribution, host, 30-32
- KondoWa2022a: distribution, host, list, 23
- KozarFoZa1996: distribution, host, 64
- Kuwana1909a: description, distribution, host, illustration, taxonomy, 161-162
- Kuwana1917a: taxonomy, 171
- LagowsGoHo2022: distribution, host, list, 11
- Laing1933: distribution, host, 676
- LincanHoCa2010: distribution, host, 6
- Lindin1914: 158
- Lindin1935a: taxonomy, 122
- LiuTa1988: description, distribution, economic importance, host, illustration, taxonomy, 63-65
- LopezMBeRe2014: endosymbionts,
- Malump2012b: distribution, 213
- Mamet1949: catalog, distribution, host, taxonomy, 12
- Mamet1952: distribution, host, 170
- Martor1976: distribution, host, 230
- Maskel1890: taxonomy, 149
- MatileEt2006: distribution, host, 177
- McKenz1960: taxonomy, 706
- MilleHeCa2016: distribution, economic importance, host, natural enemies, 148
- MillerMi2002a: taxonomy, 971
- Moghad2009: distribution, host, 18
- Moghad2013: description, distribution, illustration, structure, taxonomy, 20-21, 99
- Moghad2013a: distribution, host, 64
- Moghad2018: distribution, host, 180
- MoghadWa2022: diagnosis, distribution, illustration, key, taxonomy, 10, 42, 44-45, 144
- MohammGh2008: distribution, 155
- MonteiWoPe2019: distribution, host, 393
- Moore1988: biological control, 212
- Morris1925a: taxonomy, 487
- NurBrBe1987: distribution, life history, taxonomy, 54
- OliveiSaOl2023: DNA, phylogeny,
- PerezG2008: distribution, 216
- RenAsHu2017: DNA, phylogeny, 4, 6
- RossHaOk2012: phylogeny, taxonomy, 199
- Schmut1990: distribution, economic importance, host,
- SchmutPiKl1978: distribution, economic importance, host, 328
- SirisenSaKo2022: control, diagnosis, distribution, economic importance, host, illustration, taxonomy, 133-134
- SturzaPeNa2021: biology, distribution, host,
- Sweetm1958: biological control, economic importance, 449-458
- Takaha1932: distribution, host, taxonomy, 41
- Takaha1940a: distribution, host, 331
- Takaha1941b: distribution, host, 215
- Takaha1951a: description, distribution, host, taxonomy, 14-15
- TanakaKa2021: key, 577
- TanakaSaCh2022: key, 309
- TanakaUe2012: taxonomy, 414
- Tang1992: description, distribution, host, taxonomy, 195, 196
- Tang2001: distribution, 3
- TangHaTa1992: distribution, host, 4
- Tao1999: distribution, host, taxonomy, 13
- VanHarCoWi1990: distribution, host, 132
- Willia1931: distribution, host, life history, 113
- Willia1970DJ: description, distribution, host, illustration, taxonomy, 127-129
- Willia1985: description, distribution, host, illustration, taxonomy, 116, 117
- Willia2004a: description, distribution, host, illustration, taxonomy, 167-168
- Willia2017a: catalog, list of species, 245, 247, 250
- Willia2017b: list of species, taxonomy, 599
- WilliaBe2015: taxonomy, 163
- WilliaGr1992: description, distribution, host, illustration, taxonomy, 137-139
- WilliaMa2009a: taxonomy, 97-101
- WilliaMaSu2001: distribution, host, 73-76
- WilliaWa1988a: description, distribution, host, illustration, taxonomy, 49-51
- Wolcot1955: distribution, host, 1-5
- ZarkanApTu2021: distribution, host, 178
- ZarkanErAp2023: key, 164
- ZhangWaWa2023: key, 129
- Zimmer1948: biological control, description, distribution, host, illustration, life history, taxonomy, 177, 185-188
